Click on a red circle to go to that fossil calibration.

Time-calibrated phylogeny of ray-finned fishes, drawn vertically, with a red circle on each node calibrated by a fossil.Polypterus faraou (7 Ma)Protopsephurus luii (120.8 Ma)Polyodon tuberculata (63.1 Ma)Watsonulus eugnathoides (251.2 Ma)Anaethalion zapporum (151.2 Ma)Arratiaelops vectensis (126 Ma)Atractosteus falipoui (93.9 Ma)Baugeichthys caeruleus (129.4 Ma)Anguilla ignota (47 Ma)Serrivomer sp. (12.62 Ma)Echelus branchialis (53.7 Ma)Paralycoptera wui (107 Ma)Joffrichthys symmetropterus (58.551 Ma)Palaeonotopterus greenwoodi (93.9 Ma)Leptolepides haerteisi (150.94 Ma)Tischlingerichthys viohli (150.94 Ma)Trollichthys bolcensis (49 Ma)Eoengraulis fasoloi (49 Ma)Dorosoma petenense (1.8 Ma)Rubiesichthys gregalis (126.3 Ma)Characiformes indet. (93.9 Ma)Humboldtichthys kirschbaumi (7.246 Ma)Megapiranha paranensis (6 Ma)Lignobrycon ligniticus (24.5 Ma)Megacheirodon unicus (24.5 Ma)Salminus noriegai (7.246 Ma)Corydoras revelatus (39.5 Ma)Taubateia paraiba (24.5 Ma)Cetopangasius chaetobranchus (5.333 Ma)Astephus sp. (59.36 Ma)Ameiurus pectinatus (33.97 Ma)Pylodictis olivaris (16.3 Ma)Brachyplatystoma promagdlaena (12.8 Ma)Chrysichthys mahengeensis (45 Ma)Synodontis sp. (28.1 Ma)Amyzon aggregatum (48.88 Ma)Cyprinus maomingensis (40.14 Ma)Huashancyprinus robustispinus (23.03 Ma)Macropinna sp (7.246 Ma)Estesesox foxi (76.4 Ma)Esox kronneri (51.57 Ma)Eosalmo driftwoodensis (51.43 Ma)Hucho sp. (15.4 Ma)Oncorhynchus ('Smilodonichthys') rastrosus (8.2 Ma)Oncorhynchus keta (4.8 Ma)Paravinciguerria praecursor (93.9 Ma)Speirsaenigma lindoei (56.83 Ma)Sigmops sp. (12.62 Ma)Polypnoides laevis (41.3 Ma)Argyropelecus sp. (32.02 Ma)Argyropelecus logearti (12.62 Ma)Chauliodus testa (7.246 Ma)Chauliodus sloani (2.588 Ma)Stomias affinis (5.33 Ma)Galaxias effusus (23 Ma)Apateodus glyphodus (103.13 Ma)Alepisaurus 'ferox' (15.97 Ma)Eomyctophum koraense (32.02 Ma)Bolinichthys sp. (5.33 Ma)Homonotichthys dorsalis (93.6 Ma)Massamorichthys wilsoni (63.1 Ma)Trichophanes foliarum (33.07 Ma)Cretzeus rinaldii (69.71 Ma)Zenopsis clarus, Zenopsis tyleri, and Zenopsis hoernesi (32.02 Ma)Rhinocephalus planiceps (53.7 Ma)Nezumia lindsayi (41.3 Ma)Merluccius cf. merluccius (5.333 Ma)Gaidropsarus pilleri (13.53 Ma)Gadiculus cf. jonas (5.333 Ma)Bregmaceros filamentosus (41.3 Ma)Aipichthys velifer (98 Ma)Turkmene finitimus (54.17 Ma)Eolophotes lenis (41.3 Ma)Trachipterus mauritanicus (5.333 Ma)Stichocentrus liratus (98 Ma)Berybolcensis leptacanthus (49 Ma)Hoplopteryx lewesensis (93.6 Ma)Gephyroberyx robustus (32.02 Ma)Miobarbourisia aomori (9.83 Ma)Phyllophyarngodon longipinnis (49 Ma)Calotomus priesli (13.53 Ma)Tautoga sp. (15 Ma)Caruso brachysomus (49 Ma)Eosladenia caucasica (38 Ma)Tarkus squirei (49 Ma)Eophryne barbutii (49 Ma)Oneiroides sp. (7.42 Ma)Antennarius monodi (5.333 Ma)Ctenoplectus williamsi (53.7 Ma)Eolactoria sorbinii (49 Ma)Oligolactoria bubiki (30.28 Ma)Eospinus daniltshenkoi (54.17 Ma)Protacanthodes nimesensis (49 Ma)Carpathospinosus propheticus (26.93 Ma)Oligobalistes robustus (32.02 Ma)Balkaria histiopterygia (55.8 Ma)Austromola angerhoferi (21.12 Ma)Heptadiodon echinus (49 Ma)Archaeotetraodon winterbottomi (32.02 Ma)Eoscatophagus frontalis (49 Ma)Siganopygaeus rarus (54.17 Ma)Luvarus necopinatus (54.17 Ma)Eozanclus brevirostris (49 Ma)Proacanthurus tenuis (49 Ma)Malacanthus carosii (13.53 Ma)Lopholatilus chamaeleonticeps (13.82 Ma)Chaetodontidae indet. (Tholichthys larval stage) (29.62 Ma)Chaetodon ficheuri (5.333 Ma)Astroscopus countermani (7.246 Ma)Archoplites clarki (15.4 Ma)Gasterosteus cf. wheatlandi (13.1 Ma)Argestichthys vysotzkyi (54.17 Ma)Eocoelopoma portentosum (54.17 Ma)Eochampsodon elongatus (38 Ma)Gasterorhamphosus zuppichinii (69.71 Ma)Gerpegezhus paviai (55.8 Ma)Hippocampus samarticus (49 Ma)Carlomonnius quasigobius (49 Ma)Lepidocottus aries (23.03 Ma)Anchichanna kuldanensis (41.3 Ma)Eolates gracilis (49 Ma)Mene purdyi (55.2 Ma)Ductor vestenae (49 Ma)Oligoremora rhenana (29.62 Ma)Scomberoides spinosus (19.3 Ma)Eastmanalepes primaevus (49 Ma)Heteronectes chaneti (49 Ma)Sphyraena bolcensis (49 Ma)Eobothus minimus (49 Ma)Oligopleuronectes germanicus (29.62 Ma)Oligobothus pristinus (29.62 Ma)Eubuglossus eocenicus (41.2 Ma)Bothus sp. (11.056 Ma)Palaeopomacentrus orphae (49 Ma)Mahengechromis spp (45 Ma)Gymnogeophagus eocenicus (39.5 Ma)Nandopsis woodringi (3.6 Ma)Ramphexocoetus volans (49 Ma)Francolebias aymardi (28.1 Ma)

Table of fossil calibrations

IndexCladeFossilMin. age
1subset of PolypterusPolypterus faraou7 Ma
2Crown ChondrosteiProtopsephurus luii120.8 Ma
3Crown PolyodontidaePolyodon tuberculata63.1 Ma
4Crown HolosteiWatsonulus eugnathoides251.2 Ma
5Crown TeleosteiAnaethalion zapporum151.2 Ma
6ElopiformesArratiaelops vectensis126 Ma
7Crown LepisosteidaeAtractosteus falipoui93.9 Ma
8Albuliformes + AnguilliformesBaugeichthys caeruleus129.4 Ma
9Total group AngullidaeAnguilla ignota47 Ma
10Total group SerrivomerSerrivomer sp.12.62 Ma
11Total group EchelusEchelus branchialis53.7 Ma
12Total group OsteoglossidaeParalycoptera wui107 Ma
13Crown OsteoglossidaeJoffrichthys symmetropterus58.551 Ma
14(Mormyridae + Gymnarchidae) vs NotopteridaePalaeonotopterus greenwoodi93.9 Ma
15Total group EuteleosteiLeptolepides haerteisi150.94 Ma
16Crown OtomorphaTischlingerichthys viohli150.94 Ma
17Total group SpratteloidiniTrollichthys bolcensis49 Ma
18Crown EngraulidaeEoengraulis fasoloi49 Ma
19Crown DorosomaDorosoma petenense1.8 Ma
20Chanidae vs KneriidaeRubiesichthys gregalis126.3 Ma
21Total group CharaciformesCharaciformes indet.93.9 Ma
22MRCA Sternopygus and ApteronotusHumboldtichthys kirschbaumi7.246 Ma
23subset SerrasalmidaeMegapiranha paranensis6 Ma
24Total group LignobryconLignobrycon ligniticus24.5 Ma
25Total group SpintherobolusMegacheirodon unicus24.5 Ma
26Total group SalminusSalminus noriegai7.246 Ma
27Crown group CallichthyidaeCorydoras revelatus39.5 Ma
28Total group LoricariidaeTaubateia paraiba24.5 Ma
29Total group PangasiidaeCetopangasius chaetobranchus5.333 Ma
30Total group IctaluridaeAstephus sp.59.36 Ma
31Total group AmeiurusAmeiurus pectinatus33.97 Ma
32Total group PylodictisPylodictis olivaris16.3 Ma
33Crown BrachyplatystomaBrachyplatystoma promagdlaena12.8 Ma
34MRCA ChrysichthysChrysichthys mahengeensis45 Ma
35Total group SynodontisSynodontis sp.28.1 Ma
36Crown CatostomidaeAmyzon aggregatum48.88 Ma
37Total group CyprininiCyprinus maomingensis40.14 Ma
38Total group CyprinusHuashancyprinus robustispinus23.03 Ma
39Total group MacropinnaMacropinna sp7.246 Ma
40Total group EsocidaeEstesesox foxi76.4 Ma
41Crown EsoxEsox kronneri51.57 Ma
42Total group SalmoninaeEosalmo driftwoodensis51.43 Ma
43Total group HuchoHucho sp.15.4 Ma
44Total group OncorynchusOncorhynchus ('Smilodonichthys') rastrosus8.2 Ma
45Oncorhynchus ketaOncorhynchus keta4.8 Ma
46Total group StomiiformesParavinciguerria praecursor93.9 Ma
47Total group PlecoglossidaeSpeirsaenigma lindoei56.83 Ma
48Cylothone plus StigmopsSigmops sp.12.62 Ma
49Total group SternoptychidaePolypnoides laevis41.3 Ma
50Total group ArgyropelecusArgyropelecus sp.32.02 Ma
51Argyropelecus hemigymnusArgyropelecus logearti12.62 Ma
52Croan ChauliodusChauliodus testa7.246 Ma
53Chauliodus sloaniChauliodus sloani2.588 Ma
54Stomias affinisStomias affinis5.33 Ma
55Total group GalaxiasGalaxias effusus23 Ma
56Total group AulopiformesApateodus glyphodus103.13 Ma
57Total group AlepisaurusAlepisaurus 'ferox'15.97 Ma
58Crown MyctophiformesEomyctophum koraense32.02 Ma
59Total group BolinichthysBolinichthys sp.5.33 Ma
60Total group PolymixiiformesHomonotichthys dorsalis93.6 Ma
61Crown PercopsiformesMassamorichthys wilsoni63.1 Ma
62Amblyopsidae vs AphredoderidaeTrichophanes foliarum33.07 Ma
63Crown ZeiformesCretzeus rinaldii69.71 Ma
64Crown ZeidaeZenopsis clarus, Zenopsis tyleri, and Zenopsis hoernesi32.02 Ma
65Crown GadiformesRhinocephalus planiceps53.7 Ma
66Total group MacrouridaeNezumia lindsayi41.3 Ma
67Meluccius merlucciusMerluccius cf. merluccius5.333 Ma
68Total group GaidropsarusGaidropsarus pilleri13.53 Ma
69GadiculusGadiculus cf. jonas5.333 Ma
70Total group BregmacerosBregmaceros filamentosus41.3 Ma
71Total group LampridformesAipichthys velifer98 Ma
72Total group LamprididaeTurkmene finitimus54.17 Ma
73Total group LophotidaeEolophotes lenis41.3 Ma
74TrachipteridaeTrachipterus mauritanicus5.333 Ma
75Total group HolocentroideiStichocentrus liratus98 Ma
76Crown HolocentridaeBerybolcensis leptacanthus49 Ma
77Total group TrachichthyoideiHoplopteryx lewesensis93.6 Ma
78Total group GephyroberyxGephyroberyx robustus32.02 Ma
79Crown StephanoberyciformesMiobarbourisia aomori9.83 Ma
80Total group HypsigenyiniPhyllophyarngodon longipinnis49 Ma
81Total group CalotomusCalotomus priesli13.53 Ma
82Total group TautogaTautoga sp.15 Ma
83Crown LophiiformesCaruso brachysomus49 Ma
84MRCA Lophioides plus LophiusEosladenia caucasica38 Ma
85Total group OgcocephalidaeTarkus squirei49 Ma
86Total group AntennaridaeEophryne barbutii49 Ma
87OneiroidesOneiroides sp.7.42 Ma
88Antennarius ocellatus groupAntennarius monodi5.333 Ma
89Total group TriodontidaeCtenoplectus williamsi53.7 Ma
90Total group OstraciidaeEolactoria sorbinii49 Ma
91Crown group OstraciidaeOligolactoria bubiki30.28 Ma
92Total group BalistoideiEospinus daniltshenkoi54.17 Ma
93Total group TriacanthidaeProtacanthodes nimesensis49 Ma
94MRCA Tydemania, TriacanthodesCarpathospinosus propheticus26.93 Ma
95Crown BalistoideiOligobalistes robustus32.02 Ma
96Total group TetraodontoideiBalkaria histiopterygia55.8 Ma
97Crown MolidaeAustromola angerhoferi21.12 Ma
98Total group DiodontidaeHeptadiodon echinus49 Ma
99MRCA Sphoeroides, TetraodonArchaeotetraodon winterbottomi32.02 Ma
100Total group ScatophagidaeEoscatophagus frontalis49 Ma
101Total group SiganidaeSiganopygaeus rarus54.17 Ma
102Total group LuvarusLuvarus necopinatus54.17 Ma
103Total group ZanclusEozanclus brevirostris49 Ma
104Crown group AcanthuridaeProacanthurus tenuis49 Ma
105Total group MalacanthusMalacanthus carosii13.53 Ma
106Total group LopholatilusLopholatilus chamaeleonticeps13.82 Ma
107Total group ChaetodontidaeChaetodontidae indet. (Tholichthys larval stage)29.62 Ma
108Total group ChaetodonChaetodon ficheuri5.333 Ma
109Total group AstroscopusAstroscopus countermani7.246 Ma
110Total group ArchoplitesArchoplites clarki15.4 Ma
111Gasterosteus aculeatusGasterosteus cf. wheatlandi13.1 Ma
112Total group TrichuroideaArgestichthys vysotzkyi54.17 Ma
113Crown ScombridaeEocoelopoma portentosum54.17 Ma
114Total group ChampsodontidaeEochampsodon elongatus38 Ma
115Total group MacrorhamphosaGasterorhamphosus zuppichinii69.71 Ma
116Total group CentriscidaeGerpegezhus paviai55.8 Ma
117Total group HippocampusHippocampus samarticus49 Ma
118Total group GobiiformesCarlomonnius quasigobius49 Ma
119Total group KribiaLepidocottus aries23.03 Ma
120Total group ChannidaeAnchichanna kuldanensis41.3 Ma
121Total group LatidaeEolates gracilis49 Ma
122Total group MenidaeMene purdyi55.2 Ma
123Total group EcheneoideiDuctor vestenae49 Ma
124Total group EcheneidaeOligoremora rhenana29.62 Ma
125Total group ScomberoidiniScomberoides spinosus19.3 Ma
126Total group CaranginiEastmanalepes primaevus49 Ma
127Total group PleuronectiformesHeteronectes chaneti49 Ma
128Total group SphyraenidaeSphyraena bolcensis49 Ma
129Subset PleuronectoideiEobothus minimus49 Ma
130Subset PleuronectoideiOligopleuronectes germanicus29.62 Ma
131Total group BothidaeOligobothus pristinus29.62 Ma
132Total group SoleidaeEubuglossus eocenicus41.2 Ma
133Total group BothusBothus sp.11.056 Ma
134Total group PomacentridaePalaeopomacentrus orphae49 Ma
135Crown CichlidaeMahengechromis spp45 Ma
136Crown GeophaginiGymnogeophagus eocenicus39.5 Ma
137Total group Nandopsis haitiensisNandopsis woodringi3.6 Ma
138Total group ExocoetidaeRamphexocoetus volans49 Ma
139Total group ValenciidaeFrancolebias aymardi28.1 Ma